Dragonflies (Odonata) start their life as aquatic larvae or nymphs, passing through a series of developmental stages or ‘stadia’, and undergoing several moults as they grow. Before the final moult (emergence), metamorphosis occurs in which the larvae transform into the adult form (4). The bekko tombo has a larval period of a year, with eggs hatching from May to June, and adults emerging the following spring (2). After emergence, adults undergo a pre-reproductive phase known as the maturation period, which is when individuals normally develop their full adult colour. Odonata usually feed on flying insects and are generalised, opportunistic feeders, often congregating around abundant prey sources such as swarms of termites or near beehives (4).
There is often fierce competition between males for access to reproductive females, and females typically begin to lay eggs in water immediately after copulation, often guarded by their mate. However, females of some species can store live sperm in their body for a number of days (4).