Mrs. Appleyard's moss (Brachythecium appleyardiae)

Mrs Appleyard's moss
Loading more images and videos...

Mrs. Appleyard's moss fact file

Mrs. Appleyard's moss description


The small, rough-textured mats formed by this moss can vary in colour from a yellowish-green to pale green. Its main stems, which adhere to the underlying rock on which it grows, reach 40 mm in length. From these arise numerous crowded branches, only three to eight millimetres long. These, too, are sometimes subdivided into yet smaller branches. The slender, triangular leaves grow close to the stem, overlapping one another. They are usually flat and finely toothed or they can be un-toothed. Initially thought to be a species in the genus Eurhynchium, its capsules, with their combination of conical lids and knobbly stems led scientists to place it in the closely related Brachythecium genus as a previously undescribed species. However, it has now been proven to belong to the genus Scleropodium instead.

Also known as
Appleyard’s feather moss.
Length: up to 40 mm

Mrs. Appleyard's moss biology

Very little is known about this moss. Like all mosses, it reproduces by spores instead of seeds, although mature spore capsules have not been observed in the wild in Britain.


Mrs. Appleyard's moss range

Mrs. Appleyard's moss was thought to be endemic to Britain, and known from one site in Somerset, one in Wiltshire and six in Derbyshire. It has now also been discovered in Ireland and north-western Germany, and more new sites will probably be discovered.

You can view distribution information for this species at the National Biodiversity Network Atlas.

Mrs. Appleyard's moss habitat

This species appears to favour shaded valleys on calcareous rocks, although on one site it grows in full sunlight on a sandstone wall. In this situation, it is possible that the moss obtains its calcium from the mortar between the stones. It is usually found on dark, dry ledges that are not suitable for other species.


Mrs. Appleyard's moss status

Although previously considered a distinct species and classified as Near threatened in the UK, genetic evidence published in 2005 suggests that this moss is in fact a member of the widespread moss species, Scleropodium cespitans.


Mrs. Appleyard's moss threats

Having been found on so few sites, the status of this moss is uncertain. It is threatened by shading from un-controlled growth of ivy on one site and by secondary woodland on others. The sandstone wall on one site may require more careful maintenance to preserve the feather-moss growing on it. However, there is no direct evidence for decline. Indeed, recent evidence suggests that it could be spreading, particularly in Derbyshire. Being easily overlooked, many colonies may have been missed during surveys.


Mrs. Appleyard's moss conservation

Mrs. Appleyard's moss has been listed as a UK Biodiversity Action Plan (UKBAP) species and included in English Nature's Species Recovery Programme. It is also listed on Schedule 8 of the Wildlife and Countryside Act. However, as a member of the widespread moss species Scleropodium cespitans, as recent findings suggest, Mrs. Appleyard's moss may be removed from the BAP list, but it cannot so easily be deleted from Schedule 8.

The UK Biodiversity Action Plan for this species is available at UK BAP.
There may be further information about this species available via the National Biodiversity Network Atlas.


This information is awaiting authentication by a species expert, and will be updated as soon as possible. If you are able to help please contact:


Containing free calcium carbonate, chalky.
A species or taxonomic group that is only found in one particular country or geographic area.
Measures to conserve a species or habitat that occur outside of the natural range of the species. E.g. in zoos or botanical gardens.
Microscopic particles involved in both dispersal and reproduction. They comprise a single or group of unspecialised cells and do not contain an embryo, as do seeds.



Image credit

Mrs Appleyard's moss  
Mrs Appleyard's moss

© British Bryological Society / National Museum & Gallery, Cardiff

National Museum & Gallery, Cardiff
Cathays Park
CF10 3NP
United Kingdom


Link to this photo

Arkive species - Mrs. Appleyard's moss (Brachythecium appleyardiae) Embed this Arkive thumbnail link ("portlet") by copying and pasting the code below.

Terms of Use - The displayed portlet may be used as a link from your website to Arkive's online content for private, scientific, conservation or educational purposes only. It may NOT be used within Apps.

Read more about



MyARKive offers the scrapbook feature to signed-up members, allowing you to organize your favourite Arkive images and videos and share them with friends.

Play the Team WILD game:

Team WILD, an elite squadron of science superheroes, needs your help! Your mission: protect and conserve the planet’s species and habitats from destruction.

Conservation in Action

Which species are on the road to recovery? Find out now »

This species is featured in:

Help us share the wonders of the natural world. Donate today!


Back To Top